However, they are all underexposed a fair way.
After a few spots I switched to using spot metering on the white chests of the critters, which has netted better, but still not quite there exposure.
I was using Aperture priority mode and I'm not quite sure how to fully control spot metering in in AP.
These Cam shots are some of the better exposed. Most of the shots I can bring the exposure up well enough in PP:
Cam

PP

Cam

PP

This one was probably the best exposed 'out of the box':

So how do I get these exposures right? Guess a +EV?